Charles Darwin: The Origin of Species by means of Natural Selection

CHAPTER XII. GEOGRAPHICAL DISTRIBUTION. 4. DISPERSAL DURING THE GLACIAL PERIOD. (continued)

In the foregoing illustration, I have assumed that at the commencement of our imaginary Glacial period, the arctic productions were as uniform round the polar regions as they are at the present day. But it is also necessary to assume that many sub-arctic and some few temperate forms were the same round the world, for some of the species which now exist on the lower mountain slopes and on the plains of North America and Europe are the same; and it may be asked how I account for this degree of uniformity of the sub-arctic and temperate forms round the world, at the commencement of the real Glacial period. At the present day, the sub-arctic and northern temperate productions of the Old and New Worlds are separated from each other by the whole Atlantic Ocean and by the northern part of the Pacific. During the Glacial period, when the inhabitants of the Old and New Worlds lived further southwards than they do at present, they must have been still more completely separated from each other by wider spaces of ocean; so that it may well be asked how the same species could then or previously have entered the two continents. The explanation, I believe, lies in the nature of the climate before the commencement of the Glacial period. At this, the newer Pliocene period, the majority of the inhabitants of the world were specifically the same as now, and we have good reason to believe that the climate was warmer than at the present day. Hence, we may suppose that the organisms which now live under latitude 60 degrees, lived during the Pliocene period further north, under the Polar Circle, in latitude 66-67 degrees; and that the present arctic productions then lived on the broken land still nearer to the pole. Now, if we look at a terrestrial globe, we see under the Polar Circle that there is almost continuous land from western Europe through Siberia, to eastern America. And this continuity of the circumpolar land, with the consequent freedom under a more favourable climate for intermigration, will account for the supposed uniformity of the sub-arctic and temperate productions of the Old and New Worlds, at a period anterior to the Glacial epoch.