Charles Darwin: The Origin of Species by means of Natural Selection

CHAPTER XIII. GEOGRAPHICAL DISTRIBUTION--continued. 5. SUMMARY OF THE LAST AND PRESENT CHAPTERS.

In these chapters I have endeavoured to show that if we make due allowance for our ignorance of the full effects of changes of climate and of the level of the land, which have certainly occurred within the recent period, and of other changes which have probably occurred--if we remember how ignorant we are with respect to the many curious means of occasional transport--if we bear in mind, and this is a very important consideration, how often a species may have ranged continuously over a wide area, and then have become extinct in the intermediate tracts--the difficulty is not insuperable in believing that all the individuals of the same species, wherever found, are descended from common parents. And we are led to this conclusion, which has been arrived at by many naturalists under the designation of single centres of creation, by various general considerations, more especially from the importance of barriers of all kinds, and from the analogical distribution of subgenera, genera, and families.

With respect to distinct species belonging to the same genus, which on our theory have spread from one parent-source; if we make the same allowances as before for our ignorance, and remember that some forms of life have changed very slowly, enormous periods of time having been thus granted for their migration, the difficulties are far from insuperable; though in this case, as in that of the individuals of the same species, they are often great.

As exemplifying the effects of climatical changes on distribution, I have attempted to show how important a part the last Glacial period has played, which affected even the equatorial regions, and which, during the alternations of the cold in the north and the south, allowed the productions of opposite hemispheres to mingle, and left some of them stranded on the mountain-summits in all parts of the world. As showing how diversified are the means of occasional transport, I have discussed at some little length the means of dispersal of fresh-water productions.