Charles Darwin: The Origin of Species by means of Natural Selection

CHAPTER V. LAWS OF VARIATION.
8. SPECIFIC CHARACTERS MORE VARIABLE THAN GENERIC CHARACTERS. (continued)

On the ordinary view of each species having been independently created, why should that part of the structure, which differs from the same part in other independently created species of the same genus, be more variable than those parts which are closely alike in the several species? I do not see that any explanation can be given. But on the view that species are only strongly marked and fixed varieties, we might expect often to find them still continuing to vary in those parts of their structure which have varied within a moderately recent period, and which have thus come to differ. Or to state the case in another manner: the points in which all the species of a genus resemble each other, and in which they differ from allied genera, are called generic characters; and these characters may be attributed to inheritance from a common progenitor, for it can rarely have happened that natural selection will have modified several distinct species, fitted to more or less widely different habits, in exactly the same manner: and as these so-called generic characters have been inherited from before the period when the several species first branched off from their common progenitor, and subsequently have not varied or come to differ in any degree, or only in a slight degree, it is not probable that they should vary at the present day. On the other hand, the points in which species differ from other species of the same genus are called specific characters; and as these specific characters have varied and come to differ since the period when the species branched off from a common progenitor, it is probable that they should still often be in some degree variable--at least more variable than those parts of the organisation which have for a very long period remained constant.

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